President Thomas Wilson (Danny Glover) and other international leaders begin a secret project to ensure humanity's survival. More than 400,000 people are chosen to board "arks" that are constructed at Cho Ming, Tibet, in the southwest Chinese Himalayas. At the same time as China's People's Liberation Army (PLA) are gathering volunteers, a Buddhist monk named Nima (Osric Chau) is evacuated while his brother Tenzin (Chin Han) joins the workers in the Ark project. Additional funding for the project is raised by selling tickets to the private sector for €1 billion per person. By 2011, humanity's valuable treasures are moved to the Alps under the guise of protecting them from terrorist attacks with the help of art expert and First Daughter Dr. Laura Wilson (Thandie Newton).
In 2012, Jackson Curtis (John Cusack) is a science fiction writer in Los Angeles who works part-time as a limousine driver for the Russian billionaire, Yuri Karpov (Zlatko Burić). Jackson's ex-wife, Kate (Amanda Peet) and their children Noah (Liam James) and Lilly (Morgan Lily) live with Kate's boyfriend, plastic surgeon Gordon Silberman (Thomas McCarthy).
Jackson takes Noah and Lilly camping in Yellowstone National Park. After an encounter with Helmsley, they meet Charlie Frost (Woody Harrelson), who hosts a radio show from the park.
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